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It is possible that themecpl.dll.mui has changed, which resides in your C:\Windows\System32\en-US folder. You could right-click it and check the properties, then check the language which should say "Language: English (United States)".
sfc.exe as DC says above should probably sort that file too, if it's the culprit. Other options include a system restore which may have stamped these files since they are regarded as system, but I'm not sure. If that fails you could try copying the files from another Vista machine and overwriting (although safe mode would be best).
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