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Please take no offense, but the slowest part of either setup will be the old parts of the PC. I have used both cards, and they are really close to being on the same performance level, with the 128 MB version being a little faster every now and then. In all actuality, there are no games at the moment that really need 256 MB of video memory. Think of it this way, that would be almost half of a CD-R worth of textures that would have to be in memory all at once. It would have to be a short game, or it would have to have 10 cd's just to make a few good looking levels and characters. 128 is more than enough for the time being. But back to your PC, They would both run at the max level that your PC could feed them. So, either would be fine, and would be able to migrate into a new PC if and when you upgrade the rest.
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My computer specs? Nothing a little Windex and a paper towel won't take care of.
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